GLP-2 compounds, formulations, and uses thereof

A peptide comprising the amino acid sequence of formula I:

• His-X2-X3-Gly-X5-Phe-X7-X8-X9-X10-X11-X12-X13-X14-X15-X16-X17-X18-Ala-Arg-X21-Phe-Ile-X24-Trp-Leu-Ile-X28-Thr-Arg-Ile-Thr-X33 (formula I)
• wherein X2 is Ala, Val, or Gly;
• X3 is Asp or Glu;
• X5 is Ser or Lys;
• X7 is Ser or Lys;
• X8 is Asp, Glu, or Lys;
• X9 is Asp, Glu, or Lys;
• X10 is Met, Lys, Leu, Ile, or Nor-Leucine;
• X11 is Asn or Lys;
• X12 is Thr or Lys;
• X13 is Ile or Lys;
• X14 is Leu or Lys;
• X15 is Asp or Lys;
• X16 is Asn or Lys;
• X17 Leu or Lys;
• X18 is Ala or Lys;
• X21 is Asp or Lys;
• X24 is Asn or Lys;
• X28 is Gln or Lys;
• X33 is Asp, Glu, or Lys and at least one of X5, X7, X8, X9, X10, X11, X12, X13, X14, X15, X16, X17, and X18, X21, X24, X28, X33is a Lys.