Display substrate and display apparatus

A display substrate, comprising:

• a base substrate, comprising:
• a first opening region comprising a first opening and a first opening peripheral region surrounding the first opening;
• a display region which at least partially surrounds the first opening region and comprises:
• a first display region on a first side of the first opening region;
• and
• a second display region on a second side of the first opening region, wherein the first side and the second side are opposite to each other in a first direction, and a whole of the first display region and the second display region comprises a first pixel array;
• a second opening region, wherein the second opening region is adjacent to the first opening region, and the second opening region comprises a second opening and a second opening peripheral region surrounding the second opening;
• and
• an inter-opening region between the first opening region and the second opening region;
• a first signal line which is configured to provide a first display signal to the first pixel array, sequentially passes through the first display region, the first opening peripheral region, and the second display region along the first direction, and comprises a first lead line portion and a first winding line portion that are in the first opening peripheral region, wherein the first winding line portion partially surrounds the first opening, and the first lead line portion is connected with the first winding line portion;
• and
• a second signal line which is configured to provide a second display signal to the first pixel array, passes through the first opening peripheral region in a second direction intersecting the first direction, and comprises a second winding line portion in the first opening peripheral region, wherein the second winding line portion partially surrounds the first opening,
• wherein at least one selected from a group consisting of the inter-opening region, the first opening peripheral region, and the second opening peripheral region comprises a first virtual sub-pixel, the first virtual sub-pixel comprises a first virtual pixel circuit.