Shade tolerant lettuce

A lettuce plant comprising a modified LsKO2 gene and/or a modified Ls20ox1-B gene, wherein the homozygous presence of said genes leads to a shade tolerant phenotype,

• wherein the wild-type LsKO2 gene comprises a coding sequence having at least 70% sequence identity to SEQ ID NO.:
• 9 and the wild-type Ls20ox1-B gene comprises a coding sequence having at least 70% sequence identity to SEQ ID NO.:
• 11,
• wherein the modification comprises replacement and/or deletion and/or insertion of nucleotides resulting in an absence of functional KO2 and/or GA20ox1-B protein or wherein the modification results in the absence of the wild-type LsKO2 and/or Ls20ox1-B gene, and
• wherein the homozygous presence of one or both of the modified LsKO2 and Ls20ox1-B genes in the plant or the homozygous absence of one or both of the wild-type LsKO2 and/or Ls20ox1-B genes from the plant confers shade tolerance to the plant as compared to a plant comprising the wild-type LsKO2 and Ls20ox1-B gene and not showing shade tolerance.