Display substrate and method for operating the same, and display apparatus

A display substrate, comprising:

• a base substrate;
• and
• a display unit, provided on the base substrate and comprising a display region and a non-light-emitting region, wherein the display region comprises a sub-pixel, and the sub-pixel comprises a driving transistor and a light-emitting device;
• the driving transistor is configured to control magnitude of a driving current flowing through the light-emitting device, and comprises a gate electrode, a first pole and a second pole;
• the light-emitting device is configured to receive the driving current and be driven by the driving current to emit light, and comprises a first electrode, wherein the first electrode comprises a first portion and a second portion that are spaced apart from each other;
• the display unit further comprises:
• a connection structure which is electrically connected to the first portion of the first electrode and the second portion of the first electrode, and comprises a connection portion in the non-light-emitting region;
• and
• a first transfer electrode which is electrically connected to the first pole of the driving transistor and comprises a portion located in the non-light-emitting region, wherein the connection portion is electrically connected, in the non-light-emitting region, to the portion of the first transfer electrode located in the non-light-emitting region.