Display substrate and display panel

A display substrate, comprising:

• a base substrate;
• and
• a display region, on the base substrate and comprising a plurality of sub-pixels arranged in an array,
• wherein each of the plurality of sub-pixels comprises a light-emitting element and a pixel circuit that drives the light-emitting element to emit light, and the pixel circuit comprises a driving sub-circuit, a data writing sub-circuit, a threshold compensation sub-circuit, and a reset sub-circuit;
• the driving sub-circuit comprises a control terminal, a first terminal, and a second terminal, and is configured to control a driving current flowing through the light-emitting element;
• the data writing sub-circuit is connected to the first terminal of the driving sub-circuit, a data line, and a scan signal line, and is configured to write a data signal provided by the data line to the first terminal of the driving sub-circuit in response to a gate scan signal provided by the scan signal line;
• the threshold compensation sub-circuit is connected to the control terminal and the second terminal of the driving sub-circuit, a first voltage line, and a first reset control signal terminal, and is configured to compensate the driving sub-circuit in response to a first reset control signal received by the first reset control signal terminal and the data signal written;
• and
• the reset sub-circuit is connected to the second terminal of the driving sub-circuit, a first light-emitting control signal line, an initial signal line, and a second reset control signal terminal, and is configured to apply an initial voltage provided by the initial signal line to the second terminal of the driving sub-circuit in response to a first light-emitting control signal provided by the first light-emitting control signal line and a second reset control signal received by the second reset control signal terminal,
• wherein the reset sub-circuit comprises a first reset transistor and a second reset transistor, the threshold compensation sub-circuit comprises a threshold compensation transistor and a storage capacitor, an orthographic projection of the second reset transistor on the base substrate is between an orthographic projection of the first reset transistor on the base substrate and an orthographic projection of the threshold compensation transistor on the base substrate, and an orthographic projection of the storage capacitor on the base substrate is between the orthographic projection of the second reset transistor on the base substrate and the orthographic projection of the threshold compensation transistor on the base substrate.