O-linked glycosylation recognition motifs

A bioconjugate comprising an oligo- or polysaccharide covalently linked to a fusion protein:

• wherein the fusion protein comprises a ComP protein (ComP) glycosylation tag;
• wherein the ComP glycosylation tag comprises both a cysteine residue corresponding to the conserved cysteine residue at position 71 of SEQ ID NO:
• 2 (ComP110264:
• ENV58402.1) and a cysteine residue corresponding to the conserved cysteine residue at position 93 of SEQ ID NO:
• 2 or both a cysteine residue corresponding to the conserved cysteine residue at position 75 of SEQ ID NO:
• 1 (ComPADP1:
• AAC45886.1) and a cysteine residue corresponding to the conserved cysteine residue at position 95 of SEQ ID NO:
• 1;
• and
• wherein the fusion protein is glycosylated with the oligo- or polysaccharide on the ComP glycosylation tag at a serine residue corresponding to the conserved serine residue at position 82 of SEQ ID NO:
• 2 or position 84 of SEQ ID NO:
• 1.