Novel peptide having 4 linked ctl epitopes

A peptide having 4 linked epitopes, wherein the 4 epitope peptides are selected, optionally redundantly, from the group consisting of CTL epitope peptides:

• the peptide as shown in SEQ ID NO:
• 1 (PEP1);
• the peptide as shown in SEQ ID NO:
• 2 (PEP2);
• the peptide as shown in SEQ ID NO:
• 4 (PEP4);
• the peptide as shown in SEQ ID NO:
• 5 (PEP5);
• the peptide as shown in SEQ ID NO:
• 6 (PEP6);
• the peptide as shown in SEQ ID NO:
• 7 (PEP7);
• the peptide as shown in SEQ ID NO:
• 8 (PEP8);
• the peptide as shown in SEQ ID NO:
• 9 (PEP9);
• the peptide as shown in SEQ ID NO:
• 10 (PEP10);
• the peptide as shown in SEQ ID NO:
• 13 (PEP13);
• the peptide as shown in SEQ ID NO:
• 15 (PEP15);
• the peptide as shown in SEQ ID NO:
• 17 (PEP17);
• and the peptide as shown in SEQ ID NO:
• 18 (PEP18), linked via linkers, the peptide having 4 linked epitopes optionally comprise other peptide sequences consisting of hydrophilic amino acids, and the peptide having 4 linked epitopes has one or more features selected from the features (1) to (5) below:
• (1) the peptide comprises PEP2 at the C terminus (except for the peptide comprising PEP7 and PEP8 at the N terminus successively disposed in such order from the N terminus via a linker);
• (2) the peptide comprises PEP4 at the C terminus;
• (3) the peptide comprises PEP10 at the C terminus;
• (4) the peptide comprises PEP6 and PEP5 at the C terminus successively disposed in such order from the N terminus via a linker;
• and
• (5) the peptide comprises PEP5, PEP6, PEP9, and PEP18.