Wheat CENH3 alleles

A method of generating a haploid-inducing wheat plant, the method comprising:

• a. obtaining at least a wheat plant cell comprising an A genome, a B genome, and a D genome;
• b. mutating the B genome to obtain a homozygous knock-out mutation in a CENH3α gene;
• c. mutating the D genome to obtain a homozygous knock-out mutation in a CENH3α gene;
• d. mutating the A genome to obtain a knock-down mutation at a 5′ splice site of an intron in a CENH3α gene;
• and e. generating a wheat plant therefrom;
• whereby the wheat plant generated from step e. produces haploid progeny when crossed with a wildtype wheat plant.