Display substrate, display panel and display device

A display substrate, comprising:

• a base substrate;
• a plurality of light-emitting elements located in a display area;
• a plurality of pixel driving circuits located on the base substrate, each of the pixel driving circuits comprises an N-type transistor and a P-type transistor;
• and
• a first gate driving circuit, a second gate driving circuit and a third gate driving circuit each located on the base substrate and at least arranged in a border area at a periphery of the display area;
• the third gate driving circuit, the second gate driving circuit and the first gate driving circuit are sequentially arranged along a direction away from the display area, wherein orthographic projections of adjacent boundary areas of the first gate driving circuit and the second gate driving circuit on the base substrate are at least partially nested;
• and
• a fourth connection line extending from a region where the first gate driving circuit is located to a region where the second gate driving circuit is located, the fourth connection line being an output line of the first gate driving circuit.