Tomato plants resistant to ToBRFV, TMV, ToMV and ToMMV and corresponding resistance genes

A S. lycopersicum plant resistant to ToBRFV, comprising a mutated Tm-2-2 gene encoding a variant of the TM-2-2 protein (SEQ ID No:8),

• wherein the tyrosine (Y) at the position corresponding to position 767 of SEQ ID No:8 is not mutated or is substituted by a phenylalanine (F) or a tryptophan (W) and
• wherein said TM-2-2 protein variant comprises one of the following substitutions:
• a) the cysteine (C) at the position corresponding to position 848 of SEQ ID No:8 is substituted by an arginine (R), an asparagine or a serine is present at the position corresponding to asparagine 822 of the TM-2-2 protein (SEQ ID No:8) and a serine is present at the position corresponding to serine 825 of the TM-2-2 protein (SEQ ID No:8),
• b) the asparagine (N) at the position corresponding to position 822 of SEQ ID No:8 is substituted by a cysteine (C), a phenylalanine (F), a methionine (M), a tyrosine (Y) or a tryptophan (W), a cysteine is present at the position corresponding to cysteine 848 of the TM-2-2 protein (SEQ ID No:8) and a serine is present at the position corresponding to serine 825 of the TM-2-2 protein (SEQ ID No:8), and
• c) the serine (S) at the position corresponding to position 825 of SEQ ID No:8 is substituted by a histidine (H), a lysine (K) or a threonine (T), a cysteine is present at the position corresponding to cysteine 848 of the TM-2-2 protein (SEQ ID No:8) and an asparagine is present at the position corresponding to asparagine 822 of the TM-2-2 protein (SEQ ID No:8).