High amylose wheat with undetectable levels of SBEIIa protein

Wheat grain of Triticum aestivum comprising an embryo and starch, wherein

• (a) the embryo is homozygous for null alleles of three starch branching enzyme IIa (SBEIIa) genes,
• wherein said SBEIIa genes are an SBEIIa-A gene, an SBEIIa-B gene, and an SBEIIa-D gene,
• wherein the homozygous null alleles of at least one of the SBEIIa-A gene, the SBEIIa-B gene, and the SBEIIa-D gene comprise a null point mutation or a deletion of 1-30 basepairs, and
• (b) SBEIIa protein is undetectable in the wheat grain;
• (c) the starch comprises amylose such that the grain has an amylose content of between 50% and 90% (w/w) as a proportion of the extractable starch of the grain;
• and
• (d) the wheat grain has a germination rate of about 70% to about 100% relative to the germination rate of a wild-type wheat grain.