Acid-tolerant yeast cell, method of producing organic acid using the same, and method of producing the yeast cell

An acid-tolerant genetically engineered yeast cell comprising

• an exogenous polynucleotide that encodes lactate dehydrogenase;
• and
• a genetic modification that increases activity of an IPC synthase catalytic subunit AUR1 having at least 95% sequence identity to SEQ ID NO:
• 1;
• and optionally further comprising any one of:
• a genetic modification that increases activity of an enzyme that catalyzes introduction of a double bond to a fatty acyl site of a fatty acyl-CoA;
• a genetic modification that decreases activity of an enzyme that catalyzes formation of triacylglycerol (TG) from diacylglycerol (DG);
• or
• a combination of the genetic modifications.